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On this page you will find a list of the articles to which researchers at the Palynological laboratory have contributed.


Anderberg, Arne A. & El-Ghazaly, G. 2000. Pollen morphology in Primula sect. Carolinella (Primulaceae) and its taxonomic implications. Nordic Journal of Botany 20(1): 5-14
Cresti, M., El-Ghazaly, G. & Walles, B. 1999. Ultrastructural characteristics of Betula pendula tapetum and pollen. Grana 38(4): 194-202
El-Ghazaly, G. 2000. New and improved methods in Palynology.Plant Systematics for the 21st Century In: Plant Systematics for the 21st Century (Eds. Nordenstam, G. El-Ghazaly, M. Kassas, Portland Press, London. Pp.143-160
El-Ghazaly, G., Huysmans, S. & Smets, E.F. 2000. Pollen development of Rondeletia odorata (Rubiaceae). American Journal of Botany, 88: 166-199.
El-Ghazaly, G., Moate, R., Huysmans, S., Skvarla, J. & Rowley, J. 2000. Selected stages in pollen development in Echinodorus, Magnolia, Betula, Rondoletia, Borago and Matricaria. In: M.M. Harley, C.M. Morton and Sl Blackmore (Editors). Pollen and Spores Morphology and Biology Royal Botanic Gardens, Kew. Pp. 17-29.
Hanson, T. & El-Ghazaly, G. 2000. Development and cytochemistry of pollen and tapetum in Mitriostigma axillare (Rubiaceae). Grana 39: 65-89
Huysmans, S., El-Ghazaly, G. & Smets, E. 2000. Orbicules: still a well-hidden secret of the anther. In: Plant Systematics for the 21st Century (Eds. Nordenstam, G. El-Ghazaly, M. Kassas, Portland Press, London. Pp. 201-212
Nilson, Stefan. 2000. Fragrance glands (osmophores) in the family Oleaceae. In: Plant Systematics for the 21st Century (Eds. Nordenstam, G. El-Ghazaly, M. Kassas, Portland Press, London. Pp. 305-320
Nilsson, Siwert. 2000. Gunnar Erdtman continuous defender and innovator of palynology. In: Plant Systematics for the 21st Century (Eds. Nordenstam, G. El-Ghazaly, M. Kassas, Portland Press, London. Pp. 33-48
Nilsson, Siwert & Praglowski, J. 2000. Gunnar Errdtman a brief biographical sketch. In: Plant Systematics for the 21st Century (Eds. Nordenstam, G. El-Ghazaly, M. Kassas, Portland Press, London. Pp. 21-24
Premathilake, R., Epitawatta, S. & Nilsson, S. 1999. Pollen morphology of some selected plant species from Horton Plains, Sri Lanka. Grana 38: 289-295.


El Ghazali, G.E.B., Tsuji, S., El-Ghazaly, G.A. & Nilsson, S.: Combretaceae R. Brown. -WPSF 21(ed. S. Nilsson & G. El-Ghazaly). Scand. Univ. Press, Oslo-Stockholm-Copenhagen-Oxford-Boston. 40 pp.
El-Ghazaly, G.: Development and substructures of Pollen Grains Wall. - In: Sexual Plant Reproduction and Biotechnological applications: recent advances by molecular biology, biochemistry and morphology (ed. M. Cresti), pp. 175-200. - Springer, Wien
El-Ghazaly, G.: Tapetum and Orbicules (Ubisch bodies): Development, Morphology and Role of Pollen Grains and Tapetal Orbicules in Allergenicity. - In: Sexual Plant Reproduction and Biotechnological applications: recent advances by molecular biology, biochemistry and morphology (ed. M. Cresti), pp. 157-173. - Springer, Wien.
El-Ghazaly, G., Rowley, J. & Hesse, M.: Polarity, aperture condition and germination in pollen grains of Ephedra (Gnetales). - Plant Syst. Evol. 213: 217-231.
Huysmans, S. El-Ghazaly, G. & Smets, E.: Orbicules in Angiosperms: Morphology, Function, Distribution, and Relation with Tapetum Types. - Bot. Rev. 64(3): 240-272.
Steyn, E.M., Smith, G.F., Nilsson, S. & Grafström, E.: Pollen morphology in Aloe L. (Aloaceae). - Grana 37(1): 23-27.
Wittborn, J., Rao, K.V., El-Ghazaly, G. & Rowley, J.R.: Nanoscale similarities in the substructure of exines of Fagus pollen grains and Lycopodium spores. - Ann. Bot. 82: 141-145.


Backlund, A., Nilsson, S. Pollen morphology and the systematic position of Triplostegia (Disacales) - Taxon 46: 21-31
Ekebom, A., Nilsson, S., Saar, M. & von Hage-Hamsten, M.: A comparative study of airborne pollen concentrations of three allergenic types in Tartu (Estonia), Roma/Gotland and Stockholm (Sweden) 1990-1996. - Grana 36(6): 366-372.
El-Ghazaly, G. & Rowley, J.R.: Pollen wall of Ephedra foliata. - Palynology 21: 7-18
Gabarayeva, N., El-Ghazaly, G. Sporoderm development in Nymphaea mexicana (Nymphaeaceae). - Pl. Syst. Evol. 204: 1-19.
Gustafsson, M.H.G., Grafström, E., Nilsson, S. Pollen morphology of the Goodeniaceae and comparisions with related families - Grana 36: 185-207.
Huysmans, S., El-Ghazaly, G., Nilsson, S., Smets, E. Systematic value of tapetal orbicules: a preliminary survey of the Cinchonoideae (Rubiaceae). - Can. J. Bot. 75: 815-826.


Emilson,A., Takahashi,Y., Svensson,A., Berggren,B., and Scheynius,A. 1996. Localization of the major allergen Bet v I in birch pollen by confocal laser scanning microscopy. - Grana 35: 199-204.
Endress, M.E., Sennblad, B., Nilsson, S., Civeyrel, L., Chase, M.W., Huysmans, S., Grafström, E. and Bremer, B. 1996. A phylogenetic analysis of Apocynaceae s.str. and some related taxa in Gentianales: a multidisciplinary approach. - Opera Bot. Belg. 7: 59-102.
Henricsson, S., Westerholm, R., Nilsson, S. and Berggren, B. 1996. Chemical characterization of extractable compounds found in the coating of birch (Betula) pollen. - Grana 35: 179-184.
Isawumi, M.A, El-Ghazaly,G.and Nordenstam,B. 1996. Pollen morphology, floral microcharacters and taxonomy of the genus Baccharoides Moench (Vernonieae: Asteraceae). - Grana 35(4):204-230.
Jäger, S., Nilsson, S., Berggren, B., Pessi, A.-M., Helander, M. and Ramfjord, H. 1996. Trends of some airborne tree pollen in the Nordic countries and Austria, 1980-1993. A comparison between Stockholm, Trondheim, Turku and Vienna. - Grana 35: 171-178.
Wittborn,J., Rao,K.V.,El-Ghazaly,G. and Rowley,J.R. 1996. Substructure of spore and pollen grain exines in Lycopodium, Alnus, Betula, Fagus and Rhododendron investigated with Atomic Force an Scanning Tunnelling Microscopy. - Grana 35:185-199


Berggren, B., Nilsson, S. and Boëthius, G. 1995. Diurnal variation of airborne birch pollen in some sites in Sweden. - Grana 34: 251-259.
El-Ghazaly, G.,& Anderberg A. 1995. Pollen morphology of Phagnalon and Aliella (Asteraceae, Gnaphaleae) and its taxonomical implications. - Grana 34: 89-99.
El-Ghazaly, G., and Grafström, E. Morphological and histochemical differentiation of the pollen wall of Betula pendula Roth. during dormancy up to anthesis. - Protoplasma 187: 88-102.
El-Ghazaly, G. 1995. Pollen morphology of family Boraginaceae in Qatar. - Qatar Univ. Sci. J.-15 (19): 228-239.
El-Ghazaly, G., Takahashi, Y., Nilsson, S., Grafström, E. and Berggren, B. 1995. Orbicules in Betula pendula and their possible role in allergy. - Grana 34: 300-304.
Takahashi, Y., Nilsson, S. and Berggren, B. Aeroallergen immunoblotting with human IgE antibody. Grana 34: 357-360.


Chaudhary, R & El-Ghazaly, G. Pollen morphology of Euphorbia caputmedusae. - Grana 33:124-127.
El-Sadek, L, El-Ghazaly, G & Ayyad, M. Cytology and pollen morphology of common monocots in Mariut, Egypt I. Common species of the families Alliaceae and Liliaceae - Qatar University Science Journal.14(2):270-280.
El-Ghazaly, G., Holail, H., Sayed, O. & Aboul Dahab, O. Effect of sea level rise on Mangrove ecosystem at Al-Dhakhira - State of Qatar. - Report to UNEP. 71 pp.
El-Ghazaly, G & Kassem, M. Effect of gamma radiation on the morphology and thermophysical properties of the pollen wall of Phoenix dactylifera L..- Qatar University Science Journal.14(2):263-269.
Franzen, L.G., Hjelmroos, M., Kållberg, P., Brorström-Lunden, E., Juntto, S. & Savolainen, A-L. Yellow snow - a case study of long-range transport of dust in northern Fennoscandia, March 1991.- Atmospheric Environment Vol. 28 (22): 3587-3604.
Hjelmroos, M. & Franzen, L. Implications of recent long-distance pollen transport events on the interpretation of fossil pollen records in Fennoscandia. - Review of Palaeobotany and Palynology 82: 175-189.
Rizk, A & El-Ghazaly, G. Medicinal and poisonous plants of Qatar.- The Scientific and Applied Research Centre, University of Qatar. 270 pp.


El-Ghazaly, G. & Chaudhary, R. Pollen morphology of some species of the genus Euphorbia. - Review of Paleobotany & Palynology. 78: 293-319.
El-Ghazaly, G. & Chaudhary,R. Morphology and taxonomic application of orbicules (Ubisch bodies) in the genus Euphorbia.Grana.suppl. 2: 26-32.
El-Ghazaly, G. Larsson, K. A., Kotzamanidou-El-Ghazaly, P. & Nilsson, S. Comparison of airborne pollen grains in Stockholm and Huddinge, Sweden. - Aerobiologia 9:53-67.
Hjelmroos, M. Relationship between airborne fungal spore presence and weather variables. Cladosporium and Alternaria. - Grana 32(1): 40-47.
Hjelmroos, M., Egen, N. & Schumacher, M. Optimising the extraction of allergens in Betula pollen grains as determined by isoelectric focusing (IEF) in polyacrylamide gels. - Allergie et Immunologie 25(4): 137-140.
Hjelmroos, M. & Franzen, L.G. Implications of recent long-distance pollen transport events for the interpretation of fossil pollen records in Fennoscandia. - Review of Palaeobotany and Palynology 82: 175-189.
Hjelmroos, M. & Van Hage-Hamsten, M. Birch pollen dispersal on the Baltic Island, Gotland. - Grana, Suppl. 2: 75-79.
Nilsson, S. & Hong, D.-Y. The taxonomic significance of Aragoa-pollen (Scrophulariaceae). - Opera Bot. 121: 275-278.
Nilsson, S. Endress, M.E. & Grafström, E. On the relationship of the Apocynaceae and Periplocaceae.- Grana, Suppl. 2:3-20.


Anderberg, A. A., Karis, P.-O. & El-Ghazaly, G.: Cratystylis, an isolated genus of the Asteraceae-Cichorioideae. - Aust. Syst. Bot., 5: 81 94.
Fountain,D., Berggren,B., Nilsson,S. and Einarsson,R. 1992. Expression of birch pollen-specific IgE-binding activity in seeds and other plant parts of birch trees (Betula verrucosa Ehrh.). - Int.Arch.Allergy Immunol. 98: 370-376.
Franzen, L.G. & Hjelmroos, M. Long range transport of airborne solid particles into northern Fennoscandia - the yellow snow episode of March 1991, in the light of the Chernobyl accident of 1986. - Arctic Center publ. 4: 69-72.
Hjelmroos, M. Long distance transport of Betula pollen grains and allergic symptoms.- Aerobiologia ;8: 231-236.
Rowley, J. & El-Ghazaly, G. Lipid in wall and cytoplasm of Solidago pollen. - Grana 31: 273-283.


Ashour, M., Metwaly, A., El-Ghazaly, G. Shaksby, R. and Ali, A.: Sabkha in the State of Qatar; Geomorphological, Geographical and Biological studies, 514 pp. University of Qatar. [in Arabic, English summary].
Berglund, B.E., Hjelmroos, M. & Kolstrup, E. Vegetation and landscape through time. - In: Ecological Bulletins (Ed. Berglund, B.E.), Copenhagen, 41:109-112.
Cerceau-Larrival, M.T., Nilsson, S., Cauneau-Pigot,A., Berggren,B., Derouet, L., Verhille, A.M. and Carbonnier-Jarreau, M.C. 1991. The influence of the environment (natural and experimental) on the composition of the exine of allergenic pollen with respect to the deposition of pollutant mineral particles. - Grana 30: 532-546.
El-Ghazaly, G. Pollen Flora of Qatar. AiO Print Ltd., Odense, Denmark. 429 pp.
El-Ghazaly, G. & Nilsson, S. Development of tapetum and orbicules of Catharanthus roseus (Apocynaceae).- Systematics Association, Clarendon Press, Oxford 317-330.
El-Ghazaly, G., Yusef, H., Ahmed, M. and El-Ghazaly, P.: A study of Aerobiology of Alexandria - Egypt. - Qatar Univ. Sci. J. 11: 161-182.
Gaillard, M.-J., Berglund, B.E., Gö;ransson, H. Hjelmroos, M., Kolstrup, E. & Regnell, J. Appendix A. Chronology of the pollen diagrams from the Ystad area. - In: Ecological Bulletins (Ed. Berglund, B.E.), Copenhagen, 41:489-495.
Hjelmroos, M. Evidence of long-distance transport of Betula pollen. - Grana 30:215-228.
Hjelmroos, M. Land use in central Tunisia over the last 2000 years. - Laborativ Arkeologi 5: 15-19.
Nilsson, S. Exine/intine relation and structure in Apocynaceae pollen .- J. Palynol. (Silver Jubilee Commemoration Volume) 1990-91: 83-93.
Nilsson,S. and Berggren,B. 1991. Various methods to determine air pollutants on pollen grains. - Grana 30: 553-556.
Wallin, J.-E., Segerströ;m, L. Rosenhåll, L., Bergmann & Hjelmroos, M. Allergic symptoms caused by long-distance transported birch pollen. - Grana 30: 265-268.


Berglund, B.E., Hjelmroos, M., Bogaczewicz-Adamczak, B. & Miotk, G. The human impact in Bory Tucholskie, North Poland. A Polish-Swedish Palaeoecological research project. - Acta Univ. Lodziensis. Fol. Arch. 13: 3-11.
El-Ghareeb, R., El-Ghazaly, G., El-Sadek, L. and Nassar, M.: Morphological and Cytological studies on Polygonum in Egypt. Alexandria Univ. Sci. Bull. 140: 156-170.
El-Ghazaly, G. Development of pollen grains of Catharanthus roseus (Apocynaceae). - Review of Palaeobotany and Palynology. 64: 165-174.
El-Ghazaly, G. A comparison of pollen wall development in normal and gametocide treated plants of Triticum aestivum. - Bull.Soc.Bot.Fr., 137, Actual. bot. 2: 65-70.
El-Ghazaly, G. & Jensen, W.: Development of Wheat (Triticum aestivum) pollen wall before and after effect of a gametocide. - Canad. J. Bot. 68: 2509-2516.
Endress, M., Hesse, M., Nilsson, S., Guggisberg, A. & Zhu, J.P. The systematic position of the Holarrheninae (Apocynaceae). - Pl. Syst. Evol. 171: 157-185.
Ismail, A.M.A. and El-Ghazaly, G.: Phenological studies on Zygophyllum qatarense Hadidi from contrasting habitats. - J. Arid Env. 18: 195-205.
Nilsson, S. Taxonomic and evolutionary significance of pollen morphology in the Apocynaceae. - Pl.Syst.Evol. (Suppl.5): 91-102.