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Systematics, taxonomy, biogeography and biology of butterflies and moths (Lepidoptera) with special emphasis on the family Nepticulidae. Faunistics of Scandinavian moths fauna. Speciality: Lepidoptera (particularly Nepticulidae).
Editor of catalogue of Lepidoptera of Sweden.
Catalogus Lepidopterorum Sueciae
Svenska fjärilar
Neotropical: Costa Rica (1996); Afrotropical: The Gambia (1981, 1982); Palearctic: Cyprus (1979, 1980); Oriental: Malaysia (1991, 1992, 1994), Sri Lanka (1974), Sumatra (1992).