Hitta hit:
T-bana: Universitetet
Frescativägen 40

Ordinarie öppettider:
Tisdag-fredag 11-17
Lördag-söndag 10-18

  • Huvudmeny

Stefan Bengtson, publications


Bengtson, S., Rasmussen, B., Zi, J-W., Fletcher, I.R., Gehling, J. G.,and Bruce Runnegar B. 2021. Eocene animal trace fossils in1.7-billion-year-old metaquartzites. PNAS 2021 Vol. 118 No. 40 e2105707118. https://doi.org/10.1073/pnas.2105707118external link

Drake, H., Ivarsson, M., Heim, C., Snoeyenbos-West, O., Bengtson, S., Belivanova, V., Whitehouse, M. 2021. Fossilized anaerobic and possibly methanogenesis-fueling fungi identified deep within the Siljan impact structure. Communications Earth & Environment, DOI: 10.1038/s43247-021-00107-9external link

Ivarsson, M., Drake, M., Neubeck, A., Snoeyenbos-West, O., Belivanova, V., Bengtson, S. 2021. Introducing Paleolithobiology. GFF, DOI: 10.1080/11035897.2021.1895302external link

Little, C.T.S., Johannessen, K.C., Bengtson, S., Chan, C.S., Ivarsson, M., Slack, J.F., Broman, C., Thorseth, I.H., Grenne, T., Rouxel, O.J., Bekker, A. 2021. A late Paleoproterozoic (1.74 Ga) deep-sea, low-temperature, iron-oxidizing microbial hydrothermal vent community from Arizona, USA. Geobiology, DOI: 10.1111/gbi.12434external link.


Ivarsson, M., Drake, H., Bengtson, S., & Rasmussen, S. 2020. A cryptic alternative for the evolution of hyphae. Bioessays, 42, DOI: 10.1002/bies.201900183external link

Ivarsson, M., Drake, H., Neubeck, A., Sallstedt, T., Bengtson, S., Roberts, N.M.W & Rasmussen, B. 2020. The fossil record of igneous rock. Earth Science Reviews, 210, 103342external link.


El Albani, A., Mangano, M.G., Buatois, L.A., Bengtson, S., Riboulleau, A., Bekker, A., Konhauser, K., Lyons, T., Rollion-Bard, C., Bankole, O., Lekele Baghekema, S.G., Meunier, A., Trentesaux, A., Mazurier, A., Aubineau, J., Laforest, C., Fontaine, C., Recourt, P., Chi Fru, E., Macchiarelli, R., Reynaud, J.Y., Gauthier-Lafaye, F., and Canfield, D.E. 2019. Organism motility in an oxygenated shallow-marine environment 2.1 billion years ago. Proceedings of the National Academy of Sciences, 116(9): 3431–3436. https://doi.org/10.1073/pnas.1815721116external link

Ivarsson, M., Kilias, S.P., Broman, C., Neubeck, A., Drake, H., Chi Fru, E., Bengtson, S., Naden, J., Detsi, K., Whitehouse, M.J., 2019. Exceptional preservation of fungi as H2-bearing fluid inclusions in an early Quaternary paleo-hydrothermal system at Cape Vani, Milos, Greece. Minerals, 9: 749, doi:10.3390/min9120749external link

Yin, Z., Vargas, K., Cunningham, J., Bengtson, S., Zhu, M., Marone, F., and Donoghue, P., 2019. The early Ediacaran Caveasphaera foreshadows the evolutionary origin of animal-like embryology. Current Biology, 29(24):4307–4314.


Ivarsson, M., Bengtson, S., Drake, H., & Francis, W. , 2018. Fungi in deep subsurface environments. In: Gadd, G. Advances in Applied Microbiology, 102: 83-116, doi.org/10.1016/bs.aambs.2017.11.001external link

Ivarsson, M., Skogby, H, Phichaikamjornwut, B., Bengtson, S., Siljeström, S., Ounchanum, P., Boonsong, A., Kruachanta, M., Marone, F., Belivanova, V., Holmström, S., 2018. Intricate tunnels in garnets from soils and river sediments in Thailand – possible endolithic microborings. PLoS ONE 13(8): e0200351. doi.org/10.1371/journal.pone.0200351external link.

Sallstedt, T., Bengtson, S., Broman, C., Crill, P. M. and Canfield, D. E., 2018. Evidence of oxygenic phototrophy in ancient phosphatic stromatolites from the Paleoproterozoic Vindhyan and Aravalli Supergroups, India. Geobiology 16 (2): 139–159. DOI: 10.1111/gbi.12274external link


Bengtson, S., Sallstedt, T., Belivanova, V. & Whitehouse, M., 2017. Three-dimensional preservation of cellular and subcellular structures suggests 1.6 billion-year-old crown-group red algae. PLoS Biology, 15(3):1–38. http://journals.plos.org/plosbiology/article?id=10.1371/journal.pbio.2000735external link

Bengtson, S., Rasmussen, B., Ivarsson, M., Muhling, J., Broman, C., Marone, F., Stampanoni, M., and Bekker, A., 2017. Fungus-like mycelial fossils in 2.4 billion-year-old vesicular basalt. Nature Ecology & Evolution, online: 6 pp http://dx.doi.org/external link 10.1038/s41559-017-0141

Cunningham, J. A., Vargas,external link K., Yin,external link Z., Bengtson, S. and Donoghue, P. C. J. 2017. The Weng'an Biota (Doushantuo Formation): an Ediacaran window on soft-bodied and multicellular microorganisms. Journal of the Geological Society http://jgs.lyellcollection.org/content/early/2017/04/27/jgs2016-142external link

Davies, T. G., Rahman, I. A., Lautenschlager, S., Cunningham, J. A., Asher, R. J., Barrett, P. M., Bates, K. T., Bengtson, S. et 37 al., 2017. Open data and digital morphology.external link Proceedings of the Royal Society of London. Biological Sciences, Vol. 284, no. 1852: 1-10. Journal websiteexternal link

Doguzhaeva, L.A., Bengtson, S., Reguero, M.A. & Mörs, T., 2017. An Eocene orthocone from Antarctica shows convergent evolution of internally shelled cephalopods. PLoS ONE 12(3): e0172169. http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0172169external link

Drake, H., Ivarsson, M., Bengtson, S., Heim, C., Siljeström, S., Whitehouse, M., Broman, C., Belivanova, V., Åström, M.E. 2017. Anaerobic consortia of fungi and sulfate reducing bacteria in deep granite fractures. Nature Communications, 8:55. DOI: 10.1038/s41467-017-00094-6.external link

Ivarsson, M. och Bengtson, S. 2017. Oceanbottnarnas hemliga liv. Havsutsikt, Vol. 2: 16-18. Journal websiteexternal link

Ivarsson, M., Bengtson, S. och Carlsson, D., 2017. Är magmatiska bergarter fossilförande? Geologiskt Forum, no 94: 20-23.

Ivarsson, M., Bach, W., Broman, C., Neubeck, A., Bengtson, S. 2017. Fossilized life in subseafloor ultramafic rocks. Geomicrobiology Journal 35: 460-467.  doi.org/10.1080/01490451.2017.1370517external link.

Kouchinsky, A., Bengtson, S., Landing, E., Steiner, M., Vendrasco, M. and Ziegler, K. 2017. Terreneuvian stratigraphy and faunas from the Anabar Uplift, Siberia. Acta Palaeontologica Polonica 62 (2): 311‒440. http://www.app.pan.pl/article/item/app002892016.htmlexternal link

Yin, Z., Cunningham, J. A., Vargas, K., Bengtson, S., Zhu, M., and Donoghue, P. C. J. 2017. Nuclei and nucleoli in embryo-like fossils from the Ediacaran Weng’an Biota. Precambrian Research, 301:145–151.


Dong X-P, Vargas K, Cunningham JA, Zhang H, Liu T, Chen F, Liu J, Bengtson S, Donoghue P. C. J., 2016. Developmental biology of the early Cambrian cnidarian Olivooides. Palaeontology 59 (3): 387-407. doi: 10.1111/pala.12231external link

Ivarsson, M., Schnürer, A., Bengtson, S., Neubeck, A., 2016. Anaerobic fungi: a potential source of biological H2 in the oceanic crust. Frontiers in Microbiology 7:674, doi: 10.3389/fmicb.2016.00674.

Ivarsson, M., Bengtson, S., Neubeck, A., 2016. The igneous oceanic crust – Earth’s largest fungal habitat? Fungal Ecology 20, 249-255.


Neubeck, A., Tulej, M., Ivarsson, M., Broman, C., Riedo, A., McMahon, S., Wurz, P. & Bengtson, S., 2015. Mineralogical determination in situ of a highly heterogeneous material using a miniaturized laser ablation mass spectrometer with high spatial resolution. International Journal of Astrobiology 10.1017/S1473550415000269: 14 pp.

Donoghue, P.C.J., Cunningham, J.A., Dong, X.-P. & Bengtson, S., 2015. Embryology in deep time. In Evolutionary Developmental Biology of Invertebrates 1, edited by A. Wanninger: Springer Vienna: 45-63.

Cunningham, J.A., Vargas, K., Pengju, L., Belivanova, V., Marone, F., Martínez-Pérez, C., Guizar-Sicairos, M., Holler, M., Bengtson, S. & Donoghue, P.C.J., 2015. Critical appraisal of tubular putative eumetazoans from the Ediacaran Weng’an Doushantuo biota. Proceedings of the Royal Society. B. Biological Sciences 282(20151169): 9 pp.

Chi Fru, E., Ivarsson, M., Kilias, S.P., Frings, P.J., Hemmingsson, C., Broman, C., Bengtson, S. & Chatzitheodoridis, E., 2015. Biogenicity of an Early Quaternary iron formation, Milos Island, Greece. Geobiology online: 20 pp.

Bengtson, S. & Collins, D., 2015. Chancelloriids of the Cambrian Burgess Shale. Palaeontologia Electronica 18(1): 67 pp.

Kouchinsky, A., Bengtson, S., Clausen, S. & Vendrasco, M.J., 2015. A lower Cambrian fauna of skeletal fossils from the Emyaksin Formation, northern Siberia. Acta Palaeontologica Polonica 60(2): 421–512.


Skovsted, C.B., Kouchinsky, A., Bengtson, S. & Holmer, L.E., 2014. The problematic early Cambrian fossil Tumulduria incomperta represents the detached ventral interarea of a paterinid brachiopod. Acta Palaeontologica Polonica 59(2): 359–365.

Murdock, D.J.E., Bengtson, S., Marone, F., Greenwood, J.M. & Donoghue, P.C.J., 2014. Evaluating scenarios for the evolutionary assembly of the brachiopod body plan. Evolution and Development 16(1): 13–24.

El Albani, A., Bengtson, S., Canfield, D.E., Riboulleau, A., Rollion Bard, C., Macchiarelli, R., Ngombi Pemba, L., Hammarlund, E., Meunier, A., Moubiya Mouele, I., Benzerara, K., Bernard, S., Boulvais, P., Chaussidon, M., Cesari, C., Fontaine, C., Chi-Fru, E., Garcia Ruiz, J.M., Gauthier-Lafaye, F., Mazurier, A., Pierson-Wickmann, A.C., Rouxel, O., Trentesaux, A., Vecoli, M., Versteegh, G.J.M., White, L., Whitehouse, M. & Bekker, A., 2014. The 2.1 Ga old Francevillian biota: biogenicity, taphonomy and biodiversity. PLoS One 9(6:e99438): 18 pp.

Cunningham, J.A., Donoghue, P.C.J. & Bengtson, S., 2014. Distinguishing biology from geology in soft-tissue preservation. In Reading and Writing of the Fossil Record: Preservational Pathways to Exceptional Fossilization. The Paleontological Society Papers 20, edited by M. Laflamme, J.D. Schiffbauer & S.A.F. Darroch: 275–287.

Bengtson, S., Ivarsson, M., Astolfo, A., Belivanova, V., Broman, C., Marone, F. & Stampanoni, M., 2014. Deep-biosphere consortium of fungi and prokaryotes in Eocene sub-seafloor basalts. Geobiology 12(6): 489–496.


Ivarsson, M., Broman, C., Sturkell, E., Ormö, J., Siljeström, S., Zuilen, M.v. & Bengtson, S., 2013. Fungal colonization of an Ordovician impact-induced hydrothermal system. Scientific Reports 3(3487): 6 pp.

Ivarsson, M., Bengtson, S., Skogby, H., Belivanova, V. & Marone, F., 2013. Fungal colonies in open fractures of subseafloor basalt. Geo-Marine Letters 33(4): 233–243.

Dong, X.-p., Cunningham, J.A., Bengtson, S., Thomas, C.-W., Liu, J., Stampanoni, M. & Donoghue, P.C.J., 2013. Embryos, polyps and medusae of the early Cambrian scyphozoan Olivooides. Proceedings of the Royal Society Biological Sciences Series B 280(2130071): 1–8.

Chi Fru, E., Ivarsson, M., Kilias, S.P., Bengtson, S., Belivanova, V., Marone, F., Fortin, D., Broman, C. & Stampanoni, M., 2013. Fossilized iron bacteria reveal pathway to biological origin of banded iron formation. Nature Communications (2050).

Canfield, D.E., Ngombi Pemba, L., Hammarlund, E., Bengtson, S., Chaussidon, M., Gauthier Lafaye, F., Meunier, A., Riboulleau, A., Rollion Bard, C., Rouxel, O., Asael, D., Wickmann, A.C. & El Albani, A., 2013. Oxygen dynamics in the aftermath of the Great Oxidation of the Earth’s atmosphere. Proceedings of the National Academy of Sciences 110(42): 16736–16741.

Bengtson, S. & Ivarsson, M., 2013. Fungal symbionts in the fossil deep biosphere. In Palaeontological Association Annual Meeting. Zürich.


Murdock, D.J.E., Donoghue, P.C.J., Bengtson, S. & Marone, F., 2012. Ontogeny and microstructure of the enigmatic Cambrian tommotiid Sunnaginia Missarzhevsky 1969. Palaeontology 55(3): 661–676.

Kouchinsky, A., Bengtson, S., Runnegar, B., Skovsted, C., Steiner, M. & Vendrasco, M., 2012. Chronology of early Cambrian biomineralization. Geological Magazine 149(2): 221–251.

Ivarsson, M., Bengtson, S., Belivanova, V., Stampanoni, M., Marone, F. & Tehler, A., 2012. Fossilized fungi in subseafloor Eocene basalts. Geology 40(2): 163–166.

Huldtgren, T., Cunningham, J., Yin, C., Stampanoni, M., Marone, F., Donoghue, P.C.J. & Bengtson, S., 2012. Response to comment on “Fossilized nuclei and germination structures identify Ediacaran ‘animal embryos’ as encysting protists”. Science 335(6073): 1169-d (2 pp.).

Cunningham, J.A., Thomas, C.-W., Bengtson, S., Marone, F., Stampanoni, M., Turner, F.R., Bailey, J.V., Raff, R.A., Raff, E.C. & Donoghue, P.C.J., 2012. Experimental taphonomy of giant sulphur bacteria: implications for the interpretation of the embryo-like Ediacaran Doushantuo fossils. Proceedings of the Royal Society of London. B. Biological Sciences 279(1734): 1857–1864.

Cunningham, J.A., Thomas, C.-W., Bengtson, S., Kearns, S.L., Xiao, S., Marone, F., Stampanoni, M. & Donoghue, P.C.J., 2012. Distinguishing geology from biology in the Ediacaran Doushantuo biota relaxes constraints on the timing of the origin of bilaterians. Proceedings of the Royal Society. B. Biological Sciences 279(1737): 2369–2376.

Bengtson, S., Cunningham, J.A., Yin, C. & Donoghue, P.C.J., 2012. A merciful death for the “earliest bilaterian,” Vernanimalcula. Evolution and Development 14(5): 421–427.


Tang, F., Bengtson, S., Wang, Y., Wang, X.-l. & Yin, C.-y., 2011. Eoandromeda and the origin of Ctenophora. Evolution and Development 13(5): 408–414.

Kouchinsky, A., Bengtson, S., Clausen, S., Gubanov, A., Malinky, J.M. & Peel, J.S., 2011. A Middle Cambrian fauna of skeletal fossils from the Kuonamka Formation, northern Siberia. Alcheringa 35(1): 123–189.

Huldtgren, T., Cunningham, J., Yin, C., Stampanoni, M., Marone, F., Donoghue, P.C.J. & Bengtson, S., 2011. Fossilized nuclei and germination structures identify Ediacaran ‘animal embryos’ as encysting protists. Science 334(6063): 1696–1699.

Hammarlund, E., Canfield, D.E., Bengtson, S., Leth, P.M., Schillinger, B. & Calzada, E., 2011. The influence of sulfate concentration on soft-tissue decay and preservation. Palaeontographica Canadiana 31: 141–156.

Doguzhaeva, L. & Bengtson, S., 2011. The capsule – a newly discovered organic shell structure in the Late Cretaceous belemnite Gonioteuthis from north-west Germany. Palaeontology 54(2): 397–415.

Bengtson, S., 2011. Presentation of the 2010 Charles Schuchert Award of the Paleontological Society to Philip C. J. Donoghue. Journal of Paleontology 85(5): 1015.

Bengtson, S., 2011. Presentation of the 2010 Paleontological Society Medal to Bruce Runnegar. Journal of Paleontology 85(5): 1012.


Kouchinsky, A., Bengtson, S. & Murdock, D., 2010. A new tannuolinid problematic from the lower Cambrian of the Sukharikha River in northern Siberia. Acta Palaeontologica Polonica 55(2): 321–331.

El Albani, A., Bengtson, S., Canfield, D.E., Bekker, A., Macchiarelli, R., Mazurier, A., Hammarlund, E., Boulvais, P., Dupuy, J.-J., Fontaine, C., Fürsich, F.T., Gauthier-Lafaye, F., Janvier, P., Javaux, E., Ossa Ossa, F., Pierson-Wickmann, A.-C., Riboulleau, A., Sardini, P., Vachard, D., Whitehouse, M. & Meunier, A., 2010. Large colonial organisms with coordinated growth behaviour in oxygenated environments 2.1 billion years ago. Nature 466: 100–104.

Dong, X.-P., Bengtson, S., Gostling, N.J., Cunningham, J.A., Harvey, T.H.P., Kouchinsky, A., Repetski, J.E., Stampanoni, M. & Donoghue, P.C.J., 2010. The anatomy, taphonomy, taxonomy and systematic affinity of Markuelia: Early Cambrian to Early Ordovician scalidophorans. Palaeontology 53(6): 1291–1314.

Doguzhaeva, L.A., Mapes, R.H., Bengtson, S. & Mutvei, H., 2010. A radula and associated cephalic remains of a Late Carboniferous coleoid from Oklahoma, USA. Ferrantia 59: 37–50.

Doguzhaeva, L.A., Bengtson, S. & Mutvei, H., 2010. Structural and morphological indicators of mode of life in the Aptian lytoceratid ammonoid Eogaudryceras. In Cephalopods - Present and Past, edited by K. Tanabe, Y. Shigeta, T. Sasaki & H. Hirano. Tokyo: Tokai UP: 123–130.

Bengtson, S., 2010. A little Kraken wakes. Nature 465: 427–428.


Scott, A.C., Galtier, J., Gostling, N.J., Smith, S.Y., Collinson, M.E., Stampanoni, M., Marone, F., Donoghue, P.C.J. & Bengtson, S., 2009. Scanning electron microscopy and synchrotron radiation X-ray tomographic microscopy of 330 million year old charcoalified seed fern fertile organs. Microscopy and Microanalysis 15: 166–173.

McDowell, R., Bengtson, S. & Rasmussen, B., 2009. Tracing fossil history. Science 324: 176–177.

Kouchinsky, A., Bengtson, S., Feng, W., Kutygin, R. & Val’kov, A., 2009. The Lower Cambrian fossil anabaritids: Affinities, occurrences and systematics. Journal of Systematic Palaeontology 7(3): 241–298.

Haug, J.T., Maas, A., Waloszek, D., Donoghue, P.C.J. & Bengtson, S., 2009. A new species of Markuelia from the Middle Cambrian of Australia. Memoirs of the Association of Australasian Palaeontologists 37: 303–313.

Bengtson, S. & Rasmussen, B., 2009. New and ancient trace makers. Science 323(5912): 346–347.

Bengtson, S. & Kouchinsky, A., 2009. Virtual dissection of an early cnidarian embryo. In 9th North American Paleontological Convention. Cincinnati, Ohio.

Bengtson, S. & Collins, D., 2009. Burgess Shale chancelloriids – a prickly problem. In Walcott 2009: An International Conference on the Cambrian Explosion. Banff, Alberta.

Bengtson, S., Belivanova, V., Rasmussen, B. & Whitehouse, M., 2009. The controversial ‘Cambrian’ fossils of the Vindhyan are real but more than a billion years older. Proceedings of the National Academy of Sciences 106: 7729–7734.

Bengtson, S., 2009. Trollsländor, stenkol och syre. Rotating Beacon 2009(1): 12–13.


Tang, F., Yin, C., Bengtson, S., Liu, P., Wang, Z. & Gao, L., 2008. Octoradiate spiral organisms in the Ediacaran of South China. Acta Geologica Sinica 82(1): 27–34.

Raff, E.C., Schollaert, K.L., Nelson, D.E., Donoghue, P.C.J., Thomas, C.-W., Turner, F.R., Stein, B.D., Dong, X., Bengtson, S., Huldtgren, T., Stampanoni, M., Yin, C. & Raff, R.A., 2008. Embryo fossilization is a biological process mediated by microbial biofilms. Proceedings of the National Academy of Sciences of the United States of America 105(49): 19360–19365.

Kouchinsky, A., Bengtson, S., Gallet, Y., Korovnikov, I., Pavlov, V., Runnegar, B., Shields, G., Veizer, J., Young, E. & Ziegler, K., 2008. The SPICE carbon isotope excursion in Siberia: A combined study of the upper Middle Cambrian–lowermost Ordovician Kulyumbe River section, northwestern Siberian Platform. Geological Magazine 145(5): 609–622.

Gostling, N.J., Thomas, C.-W., Greenwood, J.M., Dong, X., Bengtson, S., Raff, E.C., Raff, R.A., Degnan, B.M., Stampanoni, M. & Donoghue, P.C.J., 2008. Deciphering the fossil record of early bilaterian embryonic development in light of experimental taphonomy. Evolution and Development 10(3): 339–349.

Chen, Z., Bengtson, S., Ming Zhou, C., Hua, H. & Yue, Z., 2008. Tube structure and original composition of Sinotubulites: shelly fossils from the late Neoproterozoic in southern Shaanxi, China. Lethaia 41(1): 37–45.

Bengtson, S., 2008. Livet på jorden. Geologiskt Forum 57: 26–29.

Bengtson, S., 2008. Krittidens flygande vinge. Rotating Beacon 2008(1): 10–12.


Kouchinsky, A., Bengtson, S., Pavlov, V., Runnegar, B., Torssander, P., Young, E. & Ziegler, K., 2007. Carbon isotope stratigraphy of the Precambrian–Cambrian Sukharikha River section, northwestern Siberian platform. Geological Magazine 144(3): 1–10.

Gostling, N.J., Donoghue, P.C.J. & Bengtson, S., 2007. The earliest fossil embryos begin to mature. Evolution and Development 9: 206–207.

Friis, E.M., Crane, P.R., Pedersen, K.R., Bengtson, S., Donoghue, P.C.J., Grimm, G.W. & Stampanoni, M., 2007. Phase-contrast X-ray microtomography links Cretaceous seeds with Gnetales and Bennettitales. Nature 450: 549–552.

Bengtson, S., Rasmussen, B. & Krapež, B., 2007. The Paleoproterozoic megascopic Stirling Biota. Paleobiology 33(3): 351–381.

Bengtson, S., 2007. Urtidens Fågel Rock. Rotating Beacon 2007(2): 5.


Tang, F., Yin, C., Bengtson, S., Liu, Y., Wang, Z., Liu, P. & Gao, L., 2006. A new discovery of macroscopic fossils from the Ediacaran Doushantuo Formation in the Yangtze Gorges area. Chinese Science Bulletin 51(12): 1487–1493.

Hagadorn, J.W., Xiao, S., Donoghue, P.C.J., Bengtson, S., Gostling, N., J., Pawlowska, M., Raff, E.C., Raff, R.A., Turner, F.R., Yin, C., Zhou, C., Yuan, X., McFeely, M.B., Stampanoni, M. & Nealson, K.H., 2006. Cellular and subcellular structure of Neoproterozoic animal embryos. Science 314: 291–294.

Donoghue, P.C.J., Kouchinsky, A., Waloszek, D., Bengtson, S., Dong, X.-p., Val’kov, A.K., Cunningham, J.A. & Repetski, J.E., 2006. Fossilized embryos are widespread but the record is temporally and taxonomically biased. Evolution and Development 8(2): 232–238.

Donoghue, P.C.J., Bengtson, S., Dong, X.-p., Gostling, N., J., Huldtgren, T., Cunningham, J.A., Yin, C., Yue, Z., Peng, F. & Stampanoni, M., 2006. Synchrotron X-ray tomographic microscopy of fossil embryos. Nature 442: 680–683.

Bengtson, S., 2006. A ghost with a bite. Nature 442(7099): 146–147.


Bengtson, S., 2005. Mineralized skeletons and early animal evolution. In Evolving Form and Function: Fossils and Development, edited by D.E.G. Briggs. New Haven, CT: Yale Peabody Museum: 101–124.

Ahlberg, P., Szaniawski, H., Clarkson, E. N. K. & Bengtson, S. 2005: Phosphatised olenid trilobites and associated fauna from the Upper Cambrian of Västergötland, Sweden. Acta Palaeontologica Polonica 50, 429-440. (PDF)

Kouchinsky, A., Bengtson, S., Pavlov, V., Val'kov, A. & Young, E. 2005. Pre-Tommotian age of the lower Pestrotsvet Formation in the Selinde section on the Siberian platform: carbon isotopic evidence. Geological Magazine 142, 319-325. (PDF)


Gubanov, A.P., Kouchinsky, A.V., Peel, J.S. & Bengtson, S., 2004. Middle Cambrian molluscan fauna of Australian aspect from northern Siberia. Alcheringa 28(1–2): 95–114.

Yin, C., Bengtson, S. & Yue, Z. 2004: Silicified and phosphatized Tianzhushania, spheroidal microfossils of possible animal origin from the Neoproterozoic of South China. Acta Palaeontologica Polonica, 49(1), 1-12. (PDF)

Bengtson, S. 2004: The body-plan explosion. Nature 430, 506. (PDF)

Rasmussen, B., Fletcher, I.R., Bengtson, S. & McNaughton, N. 2004: SHRIMP U-Pb dating of diagenetic xenotime in the Stirling Range Formation, Western Australia: 1.8 billion year minimum age for the Stirling biota. Precambrian Research 133, 329-337.   (PDF)

Bengtson, S. 2004: Early skeletal fossils. In: J. H. Lipps and B. M. Waggoner (eds.) Neoproterozoic-Cambrian Biological Revolutions. The Paleontological Society Papers 10. The Paleontological Society, pp. 67-77. (PDF)

Bengtson, S. & Budd, G. 2004: Comment on ‘Small bilaterian fossils from 40 to 55 million years before the Cambrian'. Science 306, 1291a. (PDF)


Bengtson, S., 2003. Tracing metazoan roots in the fossil record. In The New Panorama of Animal Evolution. Proceedings of the 18th International Congress of Zoology, edited by A. Legakis, S. Sfenthourakis, R. Polymeni & M. Thessalou-Legaki. Sofia: Pensoft: 289–300.

Bengtson, S. 2003: Beneath the great divide. Nature 423, 481-482. (PDF)

Warén, A., Bengtson, S., Goffredi, S.K. & Van Dover, C.L. 2003: A hot-vent gastropod with iron sulfide dermal sclerites. Science, 302, 1007. (PDF)


Rasmussen, B., Bengtson, S., Fletcher, I.R. & McNaughton, N. 2002: Discoidal impressions and trace-like fossils more than 1200 million years old. Science 296, 1112-1115. (PDF)

Kouchinsky, A. & Bengtson, S. 2002: The tube wall of Cambrian anabaritids. Acta Palaeontologica Polonica 47:3, 431-444. (PDF)

Bengtson, S. 2002: Early life, science, and wrestling. Palaeontologia Electronica 5:1external link, 3 pp.

Hammer, Ø., Bengtson, S., Malzbender, T. and Gelb, D. 2002: Imaging fossils using reflectance transformation and interactive manipulation of virtual light sources. Palaeontologia Electronica 5:1external link, 9 pp.

Bengtson, S. 2002: Tidigt liv på jorden. Geologiskt Forum, 9(35), 24-27. (PDF)

Rasmussen, B., Bengtson, S., Fletcher, I.R. & McNaughton, N. 2002: Ancient animals or something else entirely? Response. Science 298, 58-59. (PDF)

Bengtson, S. 2002. Origins and early evolution of predation. In: M. Kowalewski and P. H. Kelley (eds.) The fossil record of predation. The Paleontological Society Papers 8. The Paleontological Society, pp. 289-317. (PDF)


Kouchinsky, A., Bengtson, S., Missarzhevsky, V.V., Pelechaty, S.,Torssander, P. & Val'kov, A.K. 2001: Carbon isotope stratigraphy and the problem of a pre-Tommotian Stage in Siberia. Geological Magazine 138:4, pp. 387-396.   (PDF)

Bengtson, S. & Hou, X., 2001. The integument of Cambrian chancelloriids. Acta Palaeontologica Polonica 46(1): 1–22.

Bengtson, S. 2001: Dealing with extinct life forms. Palaeontologia Electronica 4:1external link, 5 pp.  


Bengtson, S. 2000: Let's reclaim our property. Palaeontologia Electronica 3:1 external link, 6 pp.

Bengtson, S. 2000: Teasing fossils out of shales with cameras and computers. Palaeontologia Electronica 3:1 external link, 14 pp.


Yue, Z. & Bengtson, S., 1999. Phosphatized embryo fossils from the Cambrian explosion. Chinese Science Bulletin 44(9): 842–845.

Yue, Z. & Bengtson, S., 1999. Embryonic and post-embryonic development of the Early Cambrian cnidarian Olivooides. Lethaia 32(2): 181–195.

Kouchinsky, A., Bengtson, S. & Gershwin, L.-a., 1999. Cnidarian-like embryos associated with the first shelly fossils in Siberia. Geology 27(7): 609–612.

Bengtson, S., 1999. Skeletized organisms and the evolution of skeletized tissues. In Encyclopedia of Paleontology, edited by R. Singer. Chicago, IL: Fitzroy Dearborn: 1122–1124.

Bengtson, S., 1999. L’apport spectaculaire des embryons fossilisés. La Recherche 317: 100–104.


Szaniawski, H. & Bengtson, S., 1998. Late Cambrian euconodonts from Sweden: taxonomy, evolution and stratigraphy. In Proceedings of the Sixth European Conodont Symposium, ECOS V, edited by H. Szaniawski: 7–29.

Bengtson, S. & Hou, X., 1998. The skin of chancelloriids. Palaeontological Association 42nd Annual Meeting, Portsmouth, UK: 2.

Bengtson, S., 1998. The mother of all questions. Nature 395: 560.

Bengtson, S., 1998. Animal embryos in deep time. Nature 391: 529–530.


Bengtson, S. & Yue, Z., 1997. Œufs durs. Pour la Science 241(Nov. 1997): 16–20.

Bengtson, S. & Yue, Z., 1997. Fossilized metazoan embryos from the earliest Cambrian. Science 277: 1645–1648.

Bengtson, S., Hagadorn, J., Kaesler, R. & Rumford, J., 1997. Publications in paleontology in the 21st Century. In Paleontology in the 21st Century Workshop, edited by H.R. Lane, J. Lipps, R.F. Steininger & W. Ziegler: 171–176.

Bengtson, S., 1997. New alliances for paleontology. Lethaia 30: 335–336.

Bengtson, S., 1997. Tappade jorden balansen under kambrium? Geologiskt Forum 15: 3–5.


Franzén, C. & Bengtson, S., 1996. Heinz Lowenstam’s collection of fossils. Lethaia 29: 38.

Bengtson, S., Collins, D.H. & Runnegar, B., 1996. Chancelloriid sclerite formation – turning the problem inside out. The Paleontological Society Special Publication 8: 29.

Bengtson, S., 1996. Invertebrates. In Macmillan Encyclopedia of Earth Sciences. New York, N.Y.: MacMillan: 506–509.


Franzén, C. & Bengtson, S., 1995. Manuscript processing in the digital age. Lethaia 28: 308.


Conway Morris, S. & Bengtson, S., 1994. Cambrian predators: possible evidence from boreholes. Journal of Paleontology 68(1): 1–23.

Bengtson, S. (ed.), 1994. Early Life on Earth. Nobel Symposium No. 84. New York, N.Y.: Columbia University Press.

Bengtson, S., 1994. The advent of animal skeletons. In Early Life on Earth. Nobel Symposium 84, edited by S. Bengtson. New York, N.Y.: Columbia University Press: 412–425.

Baltscheffsky, H., Bengtson, S., Bergström, J., Knoll, A.H. & Vidal, G., 1994. Preface: Approaches to understanding early life on Earth. In Early Life on Earth. Nobel Symposium 84, edited by S. Bengtson. New York, N.Y.: Columbia University Press: vii–x.


Szaniawski, H. & Bengtson, S., 1993. Origin of euconodont elements. Journal of Paleontology 67(4): 640–654.

Franzén, C. & Bengtson, S., 1993. Lethaia grows. Lethaia 26(1): 16.

Bengtson, S., 1993. Molluscan affinity of coeloscleritophorans – reply. Lethaia 26(1): 48.


Urbanek, A., Mierzejewski, P. & Bengtson, S., 1992. Fine structure of the earliest known rhabdopleurid. Lethaia 25(3): 349–350.

Towe, K.M., Bengtson, S., Fedonkin, M.A., Hofmann, H.J., Mankiewicz, C. & Runnegar, B., 1992. Described taxa of Proterozoic and selected earliest Cambrian carbonaceous remains, trace and body fossils. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge.: Cambridge U.P.: 953–1054.

Towe, K.M., Bengtson, S., Fedonkin, M.A., Hofmann, H.J., Mankiewicz, C. & Runnegar, B., 1992. Proterozoic and earliest Cambrian carbonaceous remains, trace and body fossils. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge.: Cambridge U.P.: 343–424.

Strauss, H., Bengtson, S., Myrow, P.M. & Vidal, G., 1992. Stable isotope geochemistry and palyonology of the late Precambrian to Early Cambrian sequence in Newfoundland. Canadian Journal of Earth Sciences 29(8): 1662–1673.

Hofmann, H.J., Bengtson, S., Hayes, J.M., Lipps, J.H., Schopf, J.W., Strauss, H., Summons, R.E. & Walter, M.R., 1992. Biostratigraphy and paleobiogeography of the Proterozoic. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge.: Cambridge U.P.: 487–520.

Hayes, J.M., Bengtson, S., Hofmann, H.J., Lipps, J.H., Lowe, D.R., Mankiewicz, C., Mendelson, C.V., Moore, T., Runnegar, B. & Strauss, H., 1992. Construction and use of geological, geochemical, and paleobiological databases. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge.: Cambridge U.P.: 855–864.

Bengtson, S. & Yue, Z., 1992. Predatorial borings in Late Precambrian mineralized exoskeletons. Science 257: 367–369.

Bengtson, S., Farmer, J.D., Fedonkin, M.A., Lipps, J.H. & Runnegar, B., 1992. The Proterozoic–Early Cambrian evolution of metaphytes and metazoans. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge.: Cambridge U.P.: 425–462.

Bengtson, S. & Conway Morris, S., 1992. Early radiation of biomineralizing phyla. In Origin and Early Evolution of the Metazoa, edited by J.H. Lipps & P.W. Signor. New York, N.Y.: Plenum: 447–481.

Bengtson, S., 1992. Proterozoic and earliest Cambrian skeletal metazoans. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge: Cambridge U.P.: 397–411.

Bengtson, S., 1992. The cap-shaped Cambrian fossil Maikhanella and the relationship between coeloscleritophorans and molluscs. Lethaia 25(4): 401–420.

Bengtson, S., 1992. Proterozoic and earliest Cambrian skeletal metazoans. In The Proterozoic Biosphere: A Multidisciplinary Study, edited by J.W. Schopf & C. Klein. Cambridge: Cambridge U.P.: 1017–1033.


Bengtson, S., 1991. [Review of Carter, J.H. (ed.): 1990: Skeletal biomineralization: Patterns, Processes and Evolutionary Trends.]. Nomen Nudum 20: 27–30.

Bengtson, S., 1991. Oddballs from the Cambrian start to get even. Nature 351(6323): 184–185.

Bengtson, S., 1991. The origin and extinction of phyla. Geologiska Föreningens i Stockholm Förhandlingar 113(1): 76–77.


Runnegar, B. & Bengtson, S., 1990. Origin of hard parts – early skeletal fossils. In Palaeobiology. A Synthesis, edited by D.E.G. Briggs & P.R. Crowther. Oxford: Blackwell: 24–29.

Bengtson, S., Conway Morris, S., Cooper, B.J., Jell, P.A. & Runnegar, B.N., 1990. Early Cambrian fossils from South Australia. Memoirs of the Association of Australasian Palaeontologists 9: 1–364.

Bengtson, S., 1990. Wonderful life, lovely story. Lethaia 23: 115–116.

Bengtson, S., 1990. Problematic fossil taxa. In Palaeobiology. A Synthesis, edited by D.E.G. Briggs & P.R. Crowther. Oxford: Blackwell: 442–445.

Bengtson, S., 1990. The solution to a jigsaw puzzle. Nature 345: 765–766.


Qian, Y. & Bengtson, S., 1989. Palaeontology and biostratigraphy of the Early Cambrian Meishucunian Stage in Yunnan Province, South China. Fossils and Strata 24: 1–156.

Bengtson, S., 1989. Predation in the earliest metazoan ecosystems. In Norsk Paleontologisk Forening 1969–1989, edited by H.A. Nakrem & F.-J. Lindemann: 3–4.


Bengtson, S., Fedorov, A.B., Missarzhevsky, V.V., Rozanov, A.Y., Zhegallo, E.A. & Zhuravlev, A.Y., 1987. Tumulduria incomperta and the case for Tommotian trilobites. Lethaia 20(4): 361–370.


Conway Morris, S. & Bengtson, S., 1986. The Precambrian–Cambrian boundary and geochemical perturbations. Nature 319: 696–697.

Bengtson, S. & Urbanek, A., 1986. Rhabdotubus, a Middle Cambrian rhabdopleurid hemichordate. Lethaia 19(4): 293–308.

Bengtson, S., Matthews, S.C. & Missarzhevsky, V.V., 1986. The Cambrian netlike fossil Microdictyon. In Problematic Fossil Taxa, edited by A. Hoffman & M.H. Nitecki. N.Y., & Clarendon, Oxford: Oxford U. P.: 97–115.

Bengtson, S. & Conway Morris, S., 1986. Fjälliga djur från det kambriska havet. Fauna och Flora 81(1): 27–34.

Bengtson, S., 1986. [Review of Conway Morris, S., George, J.D., Gibson, R. & Platt, H.M. (eds.) 1985: The Origins and Relationships of Lower Invertebrates.]. Palaeontological Association Circular 126 126: 11–12.

Bengtson, S., 1986. Preparing clean backgrounds in published photographic illustrations. Lethaia 19(2): 361–362.

Bengtson, S., 1986. Whittington on the Burgess Shale. Lethaia 19(2): 170.

Bengtson, S., 1986. Siliceous microfossils from the Upper Cambrian of Queensland. Alcheringa 10: 195–216.

Bengtson, S., 1986. A new Mongolian species of the Lower Cambrian genus Camenella and the problem of scleritome-based taxonomy of the Tommotiidae. Paläontologische Zeitschrift 60(1-2): 45–55.

Bengtson, S., 1986. The problem of the Problematica. In Problematic Fossil Taxa, edited by A. Hoffman & M.H. Nitecki. New York: Oxford U.P.: 3–11.


Bengtson, S., 1985. Anders Martinsson 1930–1983. Svenskt Biografiskt Lexikon 25: 224–228.

Bengtson, S., 1985. Taxonomy of disarticulated fossils. Journal of Paleontology 59: 1350–1358.

Bengtson, S., 1985. Redescription of the Lower Cambrian Halkieria obliqua Poulsen. Geologiska Föreningens i Stockholm Förhandlingar 107(2): 101–106.


Bengtson, S., Missarzhevsky, V.V. & Rozanov, A.Y., 1984. The Precambrian–Cambrian boundary: a plea for caution. Working Group on the Precambrian–Cambrian Boundary I.U.G.S.–I.G.C.P. Project No. 29 Circular–Newsletter June 1984: 14–15.

Bengtson, S. & Conway Morris, S., 1984. A comparative study of Lower Cambrian Halkieria and Middle Cambrian Wiwaxia. Lethaia 17(4): 307–329.


Bengtson, S. & Fletcher, T.P., 1983. The oldest sequence of skeletal fossils in the Lower Cambrian of southeastern Newfoundland. Canadian Journal of Earth Sciences 20(4): 525–536.

Bengtson, S., 1983. A Burgess Sale. Lethaia 16: 110.

Bengtson, S., 1983. The early history of the Conodonta. Fossils and Strata 15: 5–19.

Bengtson, S., 1983. A functional model for the conodont apparatus. Lethaia 16(1): 38.


Nilsson, R. & Bengtson, S., 1982. Problematic triactine spicules from the Upper Ordovician of Scania, Sweden. Geologiska Föreningens i Stockholm Förhandlingar 103: 377–381.


Bengtson, S. & Missarzhevsky, V.V., 1981. Coeloscleritophora – a major group of enigmatic Cambrian metazoans. In Short papers for the Second International Symposium on the Cambrian System 1981, edited by M.E. Taylor: 19–21.

Bengtson, S. & Fletcher, T.P., 1981. The succession of skeletal fossils in the basal Lower Cambrian of southeastern Newfoundland. In Short papers for the Second International Symposium on the Cambrian System 1981, edited by M.E. Taylor: 18.

Bengtson, S., 1981. The Tommotian goes international. Lethaia 14: 372.

Bengtson, S., 1981. Fossils, phylogeny, evolution and science. Lethaia 14: 166.

Bengtson, S., 1981. Fossils and evolution. Lethaia 14: 62.

Bengtson, S., 1981. En läderkorall i Gotlands silur. Fauna och Flora 76: 37–42.

Bengtson, S., 1981. Atractosella, a Silurian alcyonacean octocoral. Journal of Paleontology 55(2): 281–294.


Bengtson, S., 1980. Conodonts: the need for a functional model. Lethaia 13: 320.

Bengtson, S., 1980. Fossils and phylogeny. Lethaia 13: 276.

Bengtson, S., 1980. Redescription of the Lower Cambrian Lapworthella cornu. Geologiska Föreningens i Stockholm Förhandlingar 102(1): 53–55.


Bengtson, S., 1979. Sponges; Machaeridians; Hercolepas; Phosphatic tubes. In Lower Wenlock faunal and floral dynamics – Vattenfallet section, Gotland, edited by V. Jaanusson, S. Laufeld & R. Skoglund: 61–62; 211–212; 251.

Bengtson, S., 1978. The use and abuse of referees. Lethaia 11: 330.

Bengtson, S., 1978. Den kambriska explosionen. In Vetenskap i rörelse. Naturvetenskapliga Forskningsrådets Årsbok 1978: 35–46.

Bengtson, S., 1978. The Machaeridia – a square peg in a pentagonal hole. Thalassia Jugoslavica 12: 1–10.

Bengtson, S., 1977. Early Cambrian button-shaped phosphatic microfossils from the Siberian Platform. Palaeontology 20(4): 751–762.

Bengtson, S., 1977. Aspects of problematic fossils in the early Palaeozoic. Acta Universitatis Upsaliensis. Abstracts of Uppsala Dissertations from the Faculty of Science 415: 1–71.

Bengtson, S., Jaanusson, V. & Martinsson, A., 1976. Palaeobiology. In Acta Universitatis Upsaliensis. Uppsala University 500 Years: 85–98.

Bengtson, S., 1976. The structure of some Middle Cambrian conodonts, and the early evolution of conodont structure and function. Lethaia 9(2): 185–206.

Bengtson, S., 1973. Bibliography of the Cambrian of Norden. In Cambrian of the British Isles, Norden, and Spitsbergen, edited by C.H. Holland. London: Wiley: 258–283.

Bengtson, S., 1970. The Lower Cambrian fossil Tommotia. Lethaia 3(4): 363–392.

Bengtson, S., 1968. The problematic genus Mobergella from the Lower Cambrian of the Baltic area. Lethaia 1(4): 325–351.