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Tisdag–fredag 11–17
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Lars Werdelin, publications


Ericson, P.G.P, Irestedt, M., Zuccon, D., Larsson, P., Tison, J.-L., Emslie, S.D., Götherström, A., Hume, J.P., Werdelin, L., & Qu, Y. 2022. A 14,000-year-old genome sheds light on the evolution and extinction of a Pleistocene vulture. Communications Biology 5:857 link

Flink, T., and Werdelin, L., 2022. Digital endocasts from two late Eocene carnivores shed light on the evolution of the brain at the origin of Carnivora. Papers in Palaeontology 8:e1422. link

Hopley, P.J., Cerling, T.E., Crété, L., Werdelin, L., Mwebi, O., Manthi, F.K., & Leakey, L.N. 2022. Stable isotope analysis of carnivores from the Turkana Basin, Kenya: evidence for temporally-mixed fossil assemblages. Quaternary International. link

Jiangzuo, Q., Werdelin, L. & Sun, Y. 2022 A dwarf sabertooth cat (Felidae: Machairodontinae) from Shanxi, China, and the phylogeny of the sabertooth tribe Machairodontini. Quaternary Science Reviews 284. link

Lewis, M.E. & Werdelin, L. 2022. A revision of the genus Crocuta (Mammalia, Hyaenidae). Palaeontographica Abt. A 322(1-4): 1-115. https://10.1127/pala/2022/0120external link

Reed, K.E., Kuykendall, K.L., Herries, A.I.R., Hopley, P.J., Sponheimer, M. & Werdelin, L. 2022. Geology, fauna, and paleoenvironmental reconstructions of the Makapansgat Limeworks Australopithecus africanus-bearing paleocave. In: Reynolds, S.C. & Bobe, R. (eds) African Palaeoecology and Human Evolution. Cambridge University Press, Cambridge: 66-81.

Rowan, J., Lazagabaster, I.A., Campisano, C.J., Bibi, F., Bobe, R., Boisserie, J.-R., Frost, S.R., Getachew, T., Gilbert, C.C., Lewis, M.E., Melaku, S., Scott, E., Souron, A., Werdelin, L., Kimbel, W.H., & Reed, K.E. in press. Early Pleistocene large mammals from Maka’amitalu, Hadar, lower Awash Valley, Ethiopia. PeerJ 10:e13210. https://10.7717/peerj.13210external link


Faurby, S., Morlo, M. & Werdelin, L., 2021. CarniFOSS: A database of the body mass of fossil carnivores. Global Ecology and Biogeography. link

Flink, T., Cote, S., Rossie, J. B., Kibii, J. M. and Werdelin, L., 2021. The neurocranium of Ekweeconfractus amorui gen. et sp. nov. (Hyaenodonta, Mammalia) and the evolution of the brain in some hyaenodontan carnivores. Journal of Vertebrate Paleontology e1927748. DOI: 10.1080/02724634.2021.1927748external link

Madurell-Malapeira, J., Rodríguez-Hidalgo, A., Aouraghe, H., Haddoumi, H., Bartolini Lucenti, S., Oujaa, A., Saladié, P., Bengamra, S., Marín, J., Souhir, M., Farkouch, M., Mhamdi, H., Aissa, A.M., Werdelin, L., Chacón, M.G., Sala-Ramis, R. 2021. First small-sized Dinofelis, evidence from the Plio-Pleistocene of North Africa. Quaternary Science Reviews 265: 107028. link

Pérez-Claros, J.A., Coca-Ortega, C., & Werdelin, L. 2021. How many hyenas in North America? A quantitative perspective. Journal of Vertebrate Paleontology link

Werdelin, L. 2021. African Barbourofelinae (Mammalia, Nimravidae): A critical review. Historical Biology. link

Werdelin, L., Kitchener, A.C., Abramov, A., Veron, G. & Do Linh San, E. 2021. The scientific name of the aardwolf is Proteles cristatus. African Journal of Wildlife Research 51:149-152. link

Westbury, M.V., Barnett, R., Sandoval-Velasco, M., Gower, G., Garrett Vieira, F., de Manuel, M., Hansen, A.J., Yamaguchi, N., Werdelin, L., Marques-Bonet, T., Gilbert, M.T.P., Lorenzen, E.D. 2021. A genomic exploration of the early evolution of extant cats and their sabre-toothed relatives. Open Research Europe 1:25 link

Westbury, M.V., Le Duc, D., Duchêne, D.A., Krishnan, A., Prost, S., Rutschmann, S., Grau, J.H., Dalén, L., Weyrich, A., Norén, K., Werdelin, L., Dalerum, F., Schöneberg, T., & Hofreiter, M. 2021. Ecological specialisation and evolutionary reticulation in extant Hyaenidae. Molecular Biology and Evolution link


Faurby, S., Silvestro, D., Werdelin, L. & Antonelli, A., 2020. Brain expansion in early hominins predicts carnivore extinctions in East Africa. Ecology Letters 23: 537-544. doi: 10.1111/ele.13451external link

Kitchener, A.C., Bellemain, E., Ding, X., Kopatz, A., Kutschera, V.E., Salomashkina, V., Ruiz-García, M., Graves, T., Hou, Y., Werdelin, L. & Janke, A. 2020. Systematics, evolution and genetics of bears. In: Bears of the World: Ecology, Conservation, and Management. Penteriani, V. & Melletti, M., eds, Cambridge University Press, Cambridge, UK, 3–20.

Terhune, C.E., Curran, S., Croitor, R., Drăgușin, V., Petculescu, A., Robinson, C., Robu, M. & Werdelin, L., 2020. Early Pleistocene fauna of the Olteţ River Valley of Romania: Biochronological and biogeographic implications. Quaternary International 553, 14-33. link

Westbury, M. V., Hartmann, S., Barlow, A., Preick, M., Ridush, B., Nagel, D., Rathgeber, T., Ziegler, R., Baryshnikov, G., Sheng, G., Ludwig, G., Wiesel, I., Dalén, L., Bibi, F., Werdelin, L., Heller, R., & Hofreiter, M. 2020. Hyena palaeogenomes reveal a complex evolutionary history of cross-continental gene flow between spotted and cave hyena. Science Advances 6: eaay0456. doi: 10.1126/sciadv.aay0456external link


Lyras, G.A., Giannakopoulou, A. & Werdelin, L. 2019. The brain anatomy of an early Miocene felid from Ginn Quarry (Nebraska, USA). PalZ 93:345-355. doi: 10.1007/s12542-018-00444-9external link

Tseng, Z. J., Zazula, G. and Werdelin, L. 2019. First Fossils of Hyenas (Chasmaporthetes, Hyaenidae, Carnivora) from North of the Arctic Circle. Open Quaternary link

Werdelin, L. 2019. Comment on Thompson, J.C., Carvalho, S., Marean, C., & Alemseged, Z. Origins of the human predatory pattern. The transition to large animal exploitation by early hominins. Current Anthropology 60. DOI: 10.1086/701477external link

Werdelin L. 2019. Middle Miocene Carnivora and Hyaenodonta from Fort Ternan, western Kenya. In Bonis L. de & Werdelin L. (eds), Memorial to Stéphane Peigné: Carnivores (Hyaenodonta and Carnivora) of the Cenozoic. Geodiversitas 41(6): 267-283. link.


Adrian, B., Werdelin, L., and Grossman, A., 2018. New Miocene Carnivora (Mammalia) from Moruorot and Kalodirr, Kenya. Palaeontologia Electronica 21.1.10A 19 pp. link 

McDonald, H. G. & Werdelin, L., 2018. The sabertooth cat, Smilodon populator (Carnivora, Felidae), from Cueva del Milodón, Chile. In Werdelin, L., McDonald, H. G. & Shaw, C. A. (eds). 2018. Smilodon. The Iconic Sabertooth, 53-75.

Werdelin, L. & Flink, T., 2018. The phylogenetic context of Smilodon. In: Werdelin, L., McDonald, H. G. & Shaw, C. A. (eds). 2018. Smilodon. The Iconic Sabertooth, 14-29.

Werdelin, L., McDonald, H. G. & Shaw, C. A. (eds). 2018. Smilodon. The Iconic Sabertooth. Johns Hopkins Univeristy Press, Baltimore, 212 pp.

Villavicencio, N. A. & Werdelin, L. 2018. The Casa del Diablo cave (Puno, Peru) and the late Pleistocene demise of megafauna in the Andean Altiplano. Quaternary Science Reviews 195:21-31


Bibi, F., Pante, M., Souron, A., Stewart, K., Varela, S., Werdelin, L., Boisserie, J.-R., Fortelius, M., Hlusko, L., Njau, J. and de la Torre, I., 2017. Paleoecology of the Serengeti during the Oldowan-Acheulean transition at Olduvai Gorge, Tanzania: The mammal and fish evidence. Journal of Human Evolution. Journal websiteexternal link

Kitchener, A.C., Breitenmoser-Würsten, C., Eizirik, E., Gentry, A., Werdelin, L., Wilting, A. and Yamaguchi, N., 2017. A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group. Cat News.

Manthi, F. K., Brown, F. H., Plavcan, M. J. and Werdelin, L., 2017. Gigantic lion, Panthera leo, from the Pleistocene of Natodomeri, eastern Africa. Journal of Paleontology. Journal websiteexternal link

Viranta, S., Atickem, A., Werdelin, L. and Stenseth, N. C., 2017. Rediscovering a forgotten canid species. BMC Zoology.


Vinuesa, V., Madurell-Malapeira, J., Werdelin, L., Robles, J. M., Obradó, P., & Alba, D. M. 2016. A new skull of Hyaenictis (Carnivora, Hyaenidae) shows incipient adaptations to durophagy. Journal of Mammalian Evolution doi: 10.1007/s10914-016-9334-0

Fortelius, M., Žliobaitė, I., Kaya, F., Bibi, F., Bobe, R. Leakey, L., Leakey, M., Patterson, D., Rannikko, J. & Werdelin, L. 2016. An ecometric analysis of the fossil mammal record of the Turkana Basin. Philosophical Transactions of the Royal Society B, Biological Sciences 371: 20150232. link


Meachen, J., Dunn, R. & Werdelin, L. 2015. Carnivoran postcranial adaptations and their relationships to climate. Ecography doi: 10.1111/ecog.01656

DiMaggio, E. N., Campisano, C. J., Rowan, J., Dupont-Nivet, G., Deino, A. L., Bibi, F., Lewis, M. E., Souron, A., Garello, D., Werdelin, L., Reed, K. E., Arrowsmith, J. R. 2015. Late Pliocene Fossiliferous Sedimentary Record and the Environmental Context of early Homo from Afar, Ethiopia. Science doi: 10.1126/science.aaa1415

Werdelin, L., Lewis, M. E. & Haile-Selassie, Y. 2015. A critical review of African species of Eucyon (Mammalia; Carnivora; Canidae), with a new species from the Pliocene of the Woranso-Mille Area, Afar Region, Ethiopia. Papers in Palaeontology doi: 10.1002/spp2.1001


Chenhall, R., Martinelli, L., McLaughlin, J., Paulsen, B. S., Senior, K., Svalastog, A. L., Tunon, H., & Werdelin, L. 2014. Culture, science and bioethics - Interdisciplinary understandings of and practices in science, culture and ethics. New Zealand Online Journal of Interdisciplinary Studies Volume 1, issue 2

Barnett, R., Yamaguchi, N., Shapiro, B., Ho, S. Y. W., Barnes, I., Burger, J. Sabin, R. Werdelin, L., Cuisin, J. & Larson, G. 2014. Revealing the maternal demographic history of Panthera leo using ancient DNA and a spatially explicit genealogical analysis. BMC Evolutionary Biology 14: 70

Werdelin, L. & Wesley-Hunt, G. D. 2014. Carnivoran ecomorphology: patterns below the family level. Annales Zoologici Fennici 51: 259-268

Sheng, G.-L., Soubrier, J., Liu, J.-Y., Werdelin, L., Llamas, B., Thomson, V. A., Tuke, J., Wu, L.-W., Hou, X.-D., Chen, Q.-J., Lai, X.-L. & Cooper, A. 2014. Pleistocene cave hyenas and the recent Eurasian history of the spotted hyena, Crocuta crocuta. Molecular Ecology 23:522-533. doi: 10.1111/mec.12576

Werdelin, L., Lewis, M. E. & Haile-Selassie, Y. 2014. Mid-Pliocene Carnivora from the Woranso-Mille Area, Afar Region, Ethiopia. Journal of Mammalian Evolution. doi: 10.1007/s10914-013-9250-5


Yamaguchi, N., Driscoll, C. A., Werdelin, L., Abramov, A. A., Csorba, G., Cuisin, J., Fernholm, B., Hiermeier, M., Hills, D., Hunter, L., Itakura, H., Johansson, U. S., Kascheev, V., Krohmann, K., Martin, T., Nowak-Kemp, M., Pavlinov, I. Ya., Renoud, F., Tomsett, L., van der Mije, S., Zholnerovskaya, E., Groves, C., Kitchener, A. C., Nijman, V. & Macdonald, D. W. 2013. Locating specimens of extinct tiger (Panthera tigris) subspecies: Caspian tiger (P. t. virgata), Javan tiger (P. t. sondaica), and Balinese tiger (P. t. balica) – including previously unpublished specimens. Mammal Study 38: 187-198.

Werdelin, L. & Lewis, M. E. 2013. Temporal change in functional richness and evenness in the eastern African Plio-Pleistocene carnivoran guild. PLoS One 8(3): e57944.

Hartstone-Rose, A., Kuhn, B. F., Nalla, S., Werdelin, L. & Berger, L. R. 2013. A new species of fox from the Australopithecus sediba type locality, Malapa, South Africa. Transactions of the Royal Society of South Africa. doi:10.1080/0035919X.2012.748698

Raia, P., Carotenuto, F., Passaro, F., Piras, P., Fulgione, D., Werdelin, L., J. Saarinen, J. & Fortelius, M. 2013. Rapid action in the Palaeogene. The relationship between phenotypic and taxonomic diversification in Cenozoic mammals. Proceedings of the Royal Society B. doi: 10.1098/rspb.2012.2244


Werdelin, L. & Lewis, M. E. 2012. The taxonomic identity of the type specimen of Crocuta sivalensis (Falconer, 1867). Journal of Vertebrate Paleontology 32: 1453-1456.

Werdelin, L. & Manthi, F. K. 2012. Carnivora from the Kanapoi hominin site, northern Kenya. Journal of African Earth Sciences 64: 1-8.


Kuhn, B. F., Werdelin, L., Hartstone-Rose, A., Lacruz, R. S. & Berger, L. R. 2011. Carnivoran remains from the Malapa hominin site, South Africa. PLoS One 6(11): e26940.

Werdelin, L. 2011. A new genus and species of Felidae (Mammalia) from Rusinga Island, Kenya, with notes on early Felidae of Africa. Estudios Geológicos 67: 217-222. (Léonard Ginsburg Memorial Volume).

Andersson, K., Norman, D. & Werdelin, L. 2011. Sabretoothed carnivores and the killing of large prey. PLoS One 6(10): e24971


Hartstone-Rose, A., Werdelin, L., De Ruiter, D. J., Berger, L. R. & Churchill, S. E. 2010. The Plio-Pleistocene ancestor of wild dogs, Lycaon sekowei n. sp. Journal of Paleontology 84: 299-308.


Werdelin, L. & Simpson, S. W. 2009. The last amphicyonid in Africa. Geodiversitas 31:775-787.


Werdelin, L. & Lewis, M. E. 2008. New species of Crocuta from the Early Pliocene of Kenya, with an overview of early Pliocene hyenas of eastern Africa. Journal of Vertebrate Paleontology 28:1162-1170.

Werdelin, L. 2008. Biogeographic relationships of African carnivoran faunas 7-1.2 Ma. Comptes Rendus Palevol 7:645-656.

Dehghani, R., Wanntorp, L., Pagani, P., Källersjö, M. Werdelin, L. & Veron, G. 2008. Phylogeography of the White-tailed Mongoose (Ichneumia albicauda, Herpestidae, Carnivora, Mammalia) - a phylogenetic study based on the control region (D-loop) of mitochondrial DNA. Journal of Zoology 276: 385-393.

Turner, A., Anton, M. & Werdelin, L. 2008. Taxonomy and evolutionary patterns in the fossil Hyaenidae of Europe. Geobios 41: 677-687.

Dehghani, R. & Werdelin, L. 2008. A new small carnivoran from the Middle Miocene of Fort Ternan, Kenya. Neues Jahrbuch für Geologie und Paläontologie 248: 233-244.

Review articles, book chapters, books

Werdelin, L. & Lewis, M. E. 2013. Koobi Fora Research Project Volume 7: The Carnivora. California Academy of Sciences. x+333pp.

Werdelin, L. & Dehghani, R. 2011. Carnivora. In: Harrison, T. (ed.) Paleontology and Geology of Laetoli, Tanzania: Human Evolution in Context, Volume 2, pp. 189-232. Vertebrate Paleobiology and Paleoanthropology Series, Springer.

Lewis, M. E. & Werdelin, L. 2010. Carnivoran dispersal out of Africa during the Early Pleistocene: relevance for hominins? In: Fleagle, J. G., Shea, J. J., Grine, F. & Leakey, R. (eds.) Out of Africa 1: Who, When and Where? pp. 13-26. Vertebrate Paleobiology and Paleoanthropology Series, Springer.

Leakey, M. G. & Werdelin, L. 2010. Early Pleistocene Mammals of Africa: Background to dispersal. In: Fleagle, J. G., Shea, J. J., Grine, F. & Leakey, R. (eds.) Out of Africa 1: Who, When and Where? pp. 3-11. Vertebrate Paleobiology and Paleoanthropology Series, Springer.

Wesley-Hunt, G. D., Dehghani, R. & Werdelin, L. 2010. Comparative ecomorphology and biogeography of Herpestidae and Viverridae (Carnivora). In: Goswami, A. & Friscia, A. (eds.) Carnivoran Evolution: New Views on Phylogeny, Form, and Function, pp. 246-268. Cambridge University Press.

Werdelin, L. & Wesley-Hunt, G. D. 2010. The biogeography of carnivore ecomorphology. In: Goswami, A. & Friscia, A. (eds.) Carnivoran Evolution: New Views on Phylogeny, Form, and Function, 225-245. Cambridge University Press.

Werdelin, L., Yamaguchi, N., Johnson, W. E. & O’Brien, S. J. 2010. Phylogeny and evolution of cats. In: Macdonald, D. M. & Loveridge, A. (eds.) The Biology and Conservation of Wild Felids, pp. 59-82. Oxford University Press.

Werdelin, L. & Sanders, W. J. (eds.) 2010. Cenozoic Mammals of Africa. University of California Press. 1008 pp.

Werdelin, L. & Peigné, S. 2010. Carnivora. In: Werdelin, L. & Sanders, W. J. (eds.) Cenozoic Mammals of Africa, pp. 603-657. University of California Press.

Werdelin, L. and Cote, S. 2010. Prionogalidae (Mammalia Incertae sedis). In: Werdelin, L. & Sanders, W. J. (eds.) Cenozoic Mammals of Africa, pp. 561-562. University of California Press.

Werdelin, L. 2010. Bibymalagasia (Mammalia Incertae sedis). In: Werdelin, L. & Sanders, W. J. (eds.) Cenozoic Mammals of Africa, pp. 113-114. University of California Press.

Werdelin, L. 2010. Chronology of Neogene mammal localities. In: Werdelin, L. & Sanders, W. J. (eds.) Cenozoic Mammals of Africa, pp. 27-43. University of California Press.

Peters, C. R., Blumenschine, R. J., Hay, R. L., Livingstone, D. A., Marean, C. W., Harrison, T., Armour-Chelu, M., Andrews, P., Bernor, R. L., Bonnefille, R., & Werdelin, L. 2008. Paleoecology of the Serengeti-Mara ecosystem. In: Sinclair, A. R. E., Packer, C., Mduma, S. A. R. & Fryxell, J.M. (eds.) Serengeti III: Human impacts on ecosystem dynamics, pp. 47-94. Chicago University Press.

Popular science articles

Werdelin, L. 2013. King of beasts. Scientific American November/2013: 34-39.

Werdelin, L. 2012. Lodjurens historia. Våra Rovdjur 4/2012: 4-6.

Werdelin, L. [översättare] Dinosaurieatlas. 2008. Stockholm: BonnierCarlsen Förlag. [Translation of “Dinosaur Atlas” by John Malam, John Woodward and Michael Benton.]

Werdelin, L. [översättare] Upptäck Rovdjur. 2008. Stockholm: BonnierCarlsen Förlag. [Translation of “Insiders: Predators” by John Seidensticker and Susan Lumpkin.]