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Publications Christian Skovsted


Claybourn, T.M., Skovsted, C.B., Holmer, L.E., Pan, B., Myrow, P.M., Topper, T.P., & Brock, G.A. 2020. Brachiopods from the Byrd Group (Cambrian Series 2, Stage 4), Central Transantarctic Mountains, East Antarctica. Papers in Palaeontology. doi: 10.1002/spp2.1295external link

Li, L., Zhang, X., Skovsted, C.B., Yun, H., Pan, B. & Li, G., in press. Revisiting the molluscan fauna of the early Cambrian Xinji Formation of North China. Papers in Palaeontology. https://onlinelibrary.wiley.com/doi/full/10.1002/spp2.1289external link

Liang, Y., Holmer, L.E., Skovsted, C.B. & Zhang, Z.-F. 2020. Shell structure and affinity of the problematic early Cambrian brachiopod Heliomedusa orienta Sun and Hou, 1987. Lethaia. https://doi.org/10.1111/let.12379external link

Liu, F., Skovsted, C.B., Topper, T.P. & Zhang, Z.-F., 2020. Are hyoliths Palaeozoic lophophorates? National Science Review., 7(2): 453-469. https://academic.oup.com/nsr/advance-article/doi/10.1093/nsr/nwz161/5606729?searchresult=1external link

Liu, F., Skovsted, C.B., Topper, T.P. & Zhang, Z.-F. 2020. Revision of Triplicatella (Orthothecida, Hyolitha) with preserved digestive tracts from the early Cambrian Chengjiang Lagerstätte, South China. Historical Biology. https://doi.org/10.1080/08912963.2020.1747059external link

Luo, Q., Fariborz, G., Zhong, N., Wang, Y., Qiu, N., Skovsted, C.B., Suchý, V., Schovsbo, N.H., Morga, R., Hao, J., Liu, A., Wu, J-n., Min, X., Cao, W. & Wu, J-a., 2020. Graptolites as fossil geo-thermometers and source material of hydrocarbons: an overview of four decades of progress. Earth Science Reviews., vol. 200, article 103000. https://www.sciencedirect.com/science/article/abs/pii/S0012825219304945external link

Pan, B., Skovsted, C.B., Brock, G.A. Topper, T.P., Holmer, L.E., Li, L. & Li, G., in press. Early Cambrian organophosphatic brachiopods from the Xinji Formation, Shuiyu section, North China. Palaeoworld. https://www.sciencedirect.com/science/article/pii/S1871174X18301434external link

Skovsted, C.B., Balthasar, U., Vinther, J. & Sperling, E.A. 2020. Small shelly fossils and carbon isotopes from the early Cambrian (Stage 3-4) Mural Formation of western Laurentia. Papers in Palaeontology. https://doi.org/10.1002/spp2.1313external link

Skovsted, C.B., Martí Mus, M., Zhang, Z.-L., Pan, B., Li, L., Liu, F., Li, G. & Zhang, Z.-F. On the origin of hyolith helens. Palaeogeography, Palaeoclimatology, Palaeoecology. https://doi.org/10.1016/j.palaeo.2020.109848external link

Zhang, Z., Strotz, L.C., Topper, T.P., Chen, F., Chen, Y., Liang, Y., Zhang, Z., Skovsted, C.B. and Brock, G.A., 2020. An encrusting kleptoparasite-host interaction from the early Cambrian. Nature Communications, 11(1): 1-7.


Betts, M.J., Claybourn, T.M., Brock, G.A., Jago, J.B., Skovsted, C.B., & Paterson, J.R., 2019. Early Cambrian shelly fossils from the White Point Conglomerate, Kangaroo Island. Acta Palaeontologica Polonica 64: 489-522. https://doi.org/10.4202/app.00586.2018external link

Claybourn, T. M., Jacquet, S. M., Skovsted, C. B., Topper, T. P., Holmer, L. E. & Brock, G. A., 2019. Mollusks from the upper Shackleton Limestone (Cambrian Series 2), Central Transantarctic Mountains, East Antarctica. Journal of Paleontology 93: 437-459. https://doi.org/10.1017/jpa.2018.84
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Li, L., Zhang, X., Skovsted, C. B., Yun, H., Li, G. & Pan, B. 2019. Shell microstructures of the helcionelloid mollusc Anabarella australis from the lower Cambrian (Series 2) Xinji Formation of North China. Journal of Systematic Palaeontology. DOI: 10.1080/14772019.2018.1546236
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Pan, B., Skovsted, C.B., Sun, H. & Li, G., 2019. Early Cambrian Hyoliths from the North China Platform and their biostratigraphic and palaeogeographic significance. Alcheringa 43: 351–380. https://doi.org/10.1080/03115518.2019.1577492external link

Topper, T. P., Guo, J., Clausen, S., Skovsted, C. B. & Zhang, Z. 2019. A stem group echinoderm from the basal Cambrian of China and the origins of Ambulacraria. Nature communications 10: 1366. https://www.nature.com/articles/s41467-019-09059-3external link

Wotte, T., Skovsted, C. B., Whitehouse, M. J. & Kouchinsky, A. 2019. Isotopic evidence for well-tempered oceans during the Cambrian Explosion. Scientific Reports 9: 6330. https://doi.org/10.1038/s41598-019-42719-4external link


Betts, M.J., Paterson, J.R., Jacquet, S.M., Andrew A.S., Hall, P.A., Jago, J.B., Jagodzinski, E.A., Preiss, W.V., Crowley, J.L., Brougham, T., Mathewson, C.P., Garcia-Bellido, D.C., Topper, T.P., Skovsted, C.B. & Brock, G.A. 2018. Early Cambrian chronostratigraphy and geochronolgy of South Australia. Earth Science Reviews 185: 498-543. doi: 10.1016/j.earscirev.2018.06.005external link

Bicknell, R.R.C., Paterson, J.R., Caron, J.B. & Skovsted, C.B., 2018. The gnathobasic spine microstructure of Recent and Silurian chelicerates and the Cambrian artiopodan Sidneyia: functional and evolutionary implications. Arthropod Structure & Development. Journal websiteexternal link

Li, L., Zhang, X., Skovsted, C. B., Yun, H., Pan, B. & Li, G. 2018. Homologous shell microstructures in Cambrian hyoliths and molluscs. Palaeontology. https://onlinelibrary.wiley.com/doi/10.1111/pala.12406external link

Luo, Q., Hao, J., Skovsted, C.B., Xu, Y., Liu, Y., Wu, J., Zhang, S. & Wang, W. 2018. Optical characteristics of graptolite-bearing sediments and its implication for thermal maturity assessment. International Journal of Coal Geology 195: 386-401. doi: 10.1016/j.coal.2018.06.019external link

Pan, B, Topper, T. P., Skovsted, C. B., Miao, L. and Li, G. 2018. Occurrence of Microdictyon from the lower Cambrian Xinji Formation along the southern margin of the North China Platform. Journal of Palaeontology, Vol. 92, Special Issue 1 (Cambrian Explosion): 59-70. https://doi.org/10.1017/jpa.2017.47external link

Pan, B., Brock, G.A., Skovsted, C.B., Betts, M.J., Topper, T.P. & Li, G.-X. Paterimitra pyramidalis Laurie, 1986, the first tommotiid discovered from the early Cambrian of North China. Gondwana Research 63: 179–185.

Skovsted, C. B. & Topper, T.P. 2018. Mobergellans from the early Cambrian of Greenland and Labrador – new morphological details and implications for the functional morphology of mobergellans. Journal of Paleontology 92: 71-79.

Sperling, E.A., Balthasar, U. & Skovsted, C.B. 2018. On the edge of exceptional preservation: insights into the role of redox state in Burgess Shale-type taphonomic windows from the Mural Formation, Alberta, Canada. Emerging Topics in Life Sciences, https://doi.org/10.1042/ETLS20170163
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Zhang, Z., Skovsted, C. B. & Zhang, Z., 2018. A hyolithid without helens preserving the oldest hyolith muscle scars; palaeobiology of Paramicrocornus from the Shujingtuo Formation (Cambrian Series 2) of South China. Palaeogeography, Palaeoclimatology, Palaeoecology 489: 1-14. DOI: 10.1016/j.palaeo.2017.07.021external link


Betts, M. J., Paterson, J. R., Jago, J. B., Jacquet, S. M., Skovsted, C., Topper, T. P. and Brock, G. A. 2017, A new lower Cambrian shelly fossil biostratigraphy for South Australia:Reply, Gondwana Research Vol. 44: 262-264. Journal websiteexternal link

Betts, M. J., Paterson, J. R. Jago, J. B., Jacquet, S. M., Skovsted, C., Topper, T. P. and Brock, G. A., 2017. Global correlation of the early Cambrian of South Australia: Shelly faunaof the Dailyatia odyssei Zone. Gondwana Research, Vol. 46: 240-279. Journal websiteexternal link

Devaere, L. and Skovsted, C., 2017. New early Cambrian sclerites of Lapworthella schodakensis from NE Greenland: advancements in knowledge of lapworthellid taxonomy, sclerite growth and scleritome organization. Geological Magazine, Vol. 154 (5): 1061-1072. Journal websiteexternal link

Jacquet, S.M., Brougham, T., Skovsted, C.B., Jago, J.B., Laurie, J. R., Betts, M. J., Topper, T.P. & Brock, G. A. 2017. Watsonella crosbyi from the lower Cambrian (Terreneuvian, Stage 2) Normanville Group in South Australia. Geological Magazine 154: 1088-1104. DOI: https://doi.org/10.1017/S0016756816000704external link

Luo, Q., Hao, J., Skovsted, C., Luo, P., Khan, I., Wu, J. and Zhong, N., 2017. The organic petrology of graptolites and maturity assessment of the Wufeng–Longmaxi Formations from Chongqing, China: Insights from reflectance cross-plot analysis. International Journal of Coal Geology, Vol. 183: 161-173. Journal websiteexternal link

Skovsted, C., Knight, I., Balthasar, U. and Boyce, W. D., 2017. Depth related brachiopod faunas from the lower Cambrian Forteau Formation of southern Labrador and western Newfoundland, Canada. Palaeontologia Electronica, Vol. 20, no 3: 1-52., 54A. Journal websiteexternal link

Topper, T. P. and Skovsted, C., 2017. Keeping a lid on it: muscle scars and the mystery of the Mobergellidae. Zoological Journal of the Linnean Society, Vol. 180: 717-731

Topper, T. P., Strotz, L. C., Skovsted, C. and Holmer, L. E., 2017. Do brachiopods show substrate-related phenotypic variation? A case study from the Burgess Shale. Palaeontology, Vol. 60(2): 269-279. Journal websiteexternal link


Betts, M.J., Paterson, J.R., Jago, J.B., Jacquet, S.M., Skovsted, C.B., Topper, T.P. & Brock, G.A. 2016. A new lower Cambrian shelly fossil biostratigraphy for South Australia. Gondwana Research 36: 163-195.

Skovsted, C.B. 2016. A silicified eccentrothecimorph tommotiid from the lower Cambrian of Greenland. Bulletin of Geosciences 91: 553–559.

Skovsted, C.B., Pan, B., Topper, T.P., Betts, M.J., Li, G., & Brock, G.A., 2016. The operculum and mode of life of the lower Cambrian hyolith Cupitheca from South Australia and North China. Palaeogeography, Palaeoclimatology, Palaeoecology 443: 123-130.

Peel, J.S., Streng, M., Geyer, G., Kouchinsky, A., and Skovsted, C.B., 2016. Ovatoryctocara granulata assemblage (Cambrian Series 2–Series 3 boundary) of Løndal, North Greenland. Australasian Palaeontological Memoirs 49: 241-282.


Skovsted, C.B., Ushatinskaya, G., Holmer, L.E., Popov, L.E. & Kouchinsky, A.  2015. Taxonomy, morphology, shell structure and early ontogeny of Pelmanotreta nom. nov. from the lower Cambrian of Siberia. GFF 137: 1-8.

Skovsted, C.B., Betts, M.J., Topper, T.P. & Brock, G.A. 2015. The early Cambrian tommotiid genus Dailyatia from South Australia. Memoirs of the Association of Australasian Palaeontologists 48, 117 p.

Skovsted, C.B., Holmer, L.E., Brock, G.A., Topper, T.P. & Larsson, C.M. 2015. The early Cambrian tommotiid Kulparina rostrata from South Australia. Journal of Paleontology 89: 920-932.


Betts, M.J., Topper, T.P., Valentine, J.L., Skovsted, C.B., Paterson, J.R. & Brock, G.A. 2014. A new early Cambrian bradoriid (Arthropoda) assemblage from the northern Flinders Ranges, South Australia. Gondwana Research 25: 420-437.

Skovsted, C.B., Clausen, S., Alvaro, J.J. & Ponlevé, D. 2014. Tommotiids from the early Cambrian (Series 2, stage 3) of Morocco and the evolution of the tannuolinid scleritome and setigerous shell structures in stem group brachiopods. Palaeontology 57: 171-192.

Skovsted, C.B., Kouchinsky, A, Bengtson, S. & Holmer, L.E. 2014. The problematic Early Cambrian fossil Tumulduria incomperta represents the detached ventral interarea of a paterinid brachiopod. Acta Palaeontologica Polonica 59: 359-365.

Larsson, C.M., Skovsted, C.B., Brock, G.A., Balthasar, U., Topper, T.P. & Holmer, L.E. 2014. Paterimitra pyramidalis from South Australia; scleritome, shell structure and evolution of a lower Cambrian stem group brachiopod. Palaeontology 57: 417-446.

Topper, T.P. & Skovsted, C.B. 2014. A new name for a classic Cambrian Swedish brachiopod, Tallatella undosa (Moberg). GFF 136: 429-435.

Zhang, Z.-F., Li, G.X., Holmer, L.E., Brock, G.A., Balthasar, U., Skovsted, C.B., Fu, D.-J., Zhang, X.-L., Wang, H.-Z., Buttler, A., Zhang, Z.-L., Cao, C.-Q., Han, J., Liu, J.-N. & Shu, D.-G. 2014. An early Cambrian agglutinated tubular lophophorate with brachiopod characters. Scientific Reports 4: 4682.

Skovsted, C.B., Topper, T.P. Betts, M.J. & Brock, G.A. 2014. Associated conchs and opercula of Triplicatella disdoma (Hyolitha) from the early Cambrian of South Australia. Alcheringa 38: 138-143.


Zhang, Z., Holmer, L.E., Skovsted, C.B., Brock, G.A., Zhang, X., Shu, D., Han, J., Liu, J, Wang, H., Butler, A. And Li, G. 2013. A sclerite-bearing inferred Entoproct from the Lower Cambrian and its implications. Scientific Reports 3: 1066.

Topper, T.P., Skovsted, C.B., Harper, D.A.T. & Ahlberg, P. 2013. A bradoriid and brachiopod dominated shelly fauna from the Furongian (Cambrian) of Västergötland, Sweden. Journal of Paleontology 87: 69-83.

Topper, T.P., Holmer, L.E., Skovsted, C.B., Brock, G.A., Balthasar, U., Larsson, C.M, Pettersson  Stolk, S. & Harper, D.A.T. 2013. The oldest brachiopods from the early Cambrian of South Australia. Acta Palaeontologica Polonica 58: 93–109.

Topper, T.P., Skovsted, C.B., Peel, J.S. & Harper, D.A.T. 2013. Moulting in the lobopodian Onychodictyon from the lower Cambrian of Greenland. Lethaia 46: 490-495.


Kouchinsky, A, Bengtson, S., Runnegar, B., Skovsted, C.B., Steiner, M. & Vendrasco, M. 2012. Chronology of early Cambrian biomineralisation. Geological Magazine 149: 221-251.

Skovsted, C.B., Brock, G.A. & Topper, T.P. 2012. First occurrence of a new Ocruranus-like helcionelloid mollusc from the lower Cambrian of East Gondwana. Gondwana Research 22: 256-261.


Topper, T.P., Skovsted, C.B., Brock, G.A. & Paterson, J.R. 2011. The oldest bivalved arthropods from the early Cambrian of East Gondwana; Systematics, biostratigraphy and biogeography. Gondwana Research 19: 310-326.

Skovsted, C.B. & Peel, J.S. 2011. Hyolithellus in life position from the Lower Cambrian of North Greenland.  Journal of Paleontology 85: 37-47.

Holmer, L.E., Skovsted, C.B., Larsson, C.M, Brock, G.A. & Zhang, Z. 2011. First record of a bivalved larval shell in Early Cambrian tommotiids and its phylogenetic significance. Palaeontology 54: 235-239.

Skovsted, C.B., Brock, G.A., Topper, T.P., Paterson, J.R. & Holmer, L.E. 2011. Scleritome construction, biofacies, biostratigraphy and systematics of the tommotiid Eccentrotheca helenia sp. nov. from the early Cambrian of South Australia. Palaeontology 54: 253-286.

Topper, T.P., Brock, G.A., Skovsted, C.B. & Paterson, J.R. 2011. Microdictyon plates from the lower Cambrian Ajax Limestone of South Australia: Implications for species taxonomy and diversity. Alcheringa 35: 427-443.

Skovsted, C.B., Brock, G.A. & Topper, T.P. 2011. Sclerite fusion in the problematic early Cambrian spine-like fossil Stoibostrombus from South Australia. Bulletin of Geosciences 86: 651–658.

Holmer, L.E., Skovsted, C.B., Brock, G.A. & Popov, L.E. 2011. An early Cambrian chileate brachiopod from South Australia and its phylogenetic significance. Memoirs of the Association of Australasian Palaeontologists 41: 289-294.


Popov, L.E., Bassett, M.G., Holmer, L.E. Skovsted, C.B. & Zuykov, M. 2010. Earliest Ontogeny Of Early Palaeozoic Craniiformea: Implications For Brachiopod Phylogeny. Lethaia 43: 323-333.

Topper, T.P., Brock, G.A., Skovsted, C.B. & Paterson, J.R. 2010. Palaeoscolecid Scleritome Fragments With Hadimopanella Plates From The Early Cambrian Of South Australia. Geological Magazine 147: 86-97.

Skovsted, C.B. & Peel, J.S. 2010. Early Cambrian Brachiopods and other Small Shelly Fossils From The Basal Kinzers Formation Of Pennsylvania. Journal of Paleontology 84: 754-762.

Skovsted, C.B., Holmer, L.E., Streng, M. & Knight, I. 2010. Setatella significans, a new name for mickwitziid stem group brachiopods from the lower Cambrian of Greenland and Labrador. GFF 132: 117-122.

Hu, S, Zhang, Z., Holmer, L.E. & Skovsted, C.B. 2010. First record of soft-part preservation in a linguliform brachiopod from the Lower Cambrian Wulongqing Formation (Guanshan Fauna) of Yunnan, South China. Acta Palaeontologica Polonica 55: 495-505.